Mean number of species (i.e., species richness) and individuals (i.e., abundance) of native and nonnative beetle species collected in port areas (“Port”) vs. surrounding natural areas (“Outside”). Trends are shown separately for the different beetle families. Abundance is log-transformed according to data transformation used in statistical analysis. P-values: * = 0.01 - 0.05; ** = 0.01 - 0.001; *** = < 0.001.