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  • Article title
  • Abstract
  • Keywords
  • Introduction
  • Materials and methods
    • Study sites
    • Study species
    • Two-factor field exclosure experiment in steppe and forest
    • Statistical analysis
  • Results
  • Discussion
    • Seed limitation, disturbance, and naturalizations
    • Competition in PNW coniferous forests
    • Conclusions and future directions
  • Acknowledgments
  • References
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