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  • Article title
  • Abstract
  • Key words
  • Introduction
  • Methods
    • Study site and materials
    • Experiment design
    • Root trait measurement
    • Statistical analyses
  • Results
    • Trait variation between ranges
    • Effect of soil nutrients on root traits
    • Trait covariation
  • Discussion
    • Variation in root traits between ranges
    • Response to soil nutrients
    • Changes in trait syndromes between ranges
  • Conclusion
  • Additional information
    • Conflict of interest
    • Ethical statement
    • Funding
    • Author contributions
    • Author ORCIDs
    • Data availability
  • References
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