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  • Article title
  • Abstract
  • Key words
  • Introduction
  • Concept for integrating fine-scale environmental heterogeneity and functional changes into spatial models of invader-ecosystem interactions
  • Environmental context
  • Structure and function
  • Functional tracers of invader impact
  • Integration: spatial modelling of functional changes and impact assessment across different stages of invasion
  • Concluding remarks
  • Acknowledgements
  • Additional information
    • Conflict of interest
    • Ethical statement
    • Funding
    • Author contributions
    • Author ORCIDs
    • Data availability
  • References
  • Supplementary materials
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