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  • ContentsContents
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  • Article title
  • Abstract
  • Key words
  • Introduction
  • Materials and methods
    • Collection of specimens
    • Terminology of invasion: the case of land flatworms
    • Data on geographical distribution
    • Molecular barcoding and population genetics
    • DNA extraction
    • Genetic characterization
    • Intraspecific genetic variation
    • Data resources
  • Results
    • Temporal and spatial distribution of non-native land flatworms in the Netherlands
    • Species of Geoplaninae
    • Species of Bipaliinae
    • Temporal distribution in Europe
    • Species of Bipaliinae
    • Species of Geoplaninae
    • Barcoding results and intraspecific genetic variation
  • Discussion
    • The Netherlands: gateway of horticulture to the rest of Europe
    • Negative impact of the particular non-native land flatworms in this study
    • Bipalium kewense: from indoors to outdoors
    • Rapid rise of O. nungara and the first report of O. anthropophila in Europe
    • Genetic variation and origin of Obama nungara in Europe
    • Importance of citizen science and museum collections
    • Challenges in data collection
    • Importance of habitat definitions
  • Conclusion
    • Use of photos, icons and personal information
  • Acknowledgements
  • Additional information
    • Conflict of interest
    • Ethical statement
    • Funding
    • Author contributions
    • Author ORCIDs
    • Data availability
  • References
  • Supplementary materials
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