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  • Article title
  • Abstract
  • Key words
  • Introduction
  • Material and methods
    • Study area and sampling
    • Environmental characterisation
    • Trace element analysis
    • Stable isotope analysis of crayfish
    • Behavioural tests
    • Epibiotic associate analysis
    • Data analysis
  • Results
    • Environmental characterisation
    • Elemental analysis
    • Stable isotope analysis of signal crayfish
    • Behavioural tests
    • Epibiotic associate analysis
  • Discussion
    • Elemental accumulation
    • Possible mechanisms explaining different elemental accumulation along the invasion gradient
    • Management implications
  • Conclusion
  • Additional information
    • Conflict of interest
    • Ethical statement
    • Funding
    • Author contributions
    • Author ORCIDs
    • Data availability
  • References
  • Supplementary material
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