Effects of invasiveness of the invasive species ( A) and abundances of the natives (B) on differences between invasive and native species, respectively. S, M and W indicate that strongly, moderately and weakly invasive species are compared with their co-occurring natives, respectively. D, C and R indicate that widespread dominant, common and rare natives are compared with their co-occurring invasives, respectively. The traits whose interspecific differences were not affected by those factors were not shown. See Figure 1 for trait abbreviations. The figures between brackets on the left indicate the numbers of species pairs included in the analyses.