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  • Article title
  • Abstract
  • Keywords
  • Introduction
  • Materials and methods
    • Study species
    • Seed collection and preparation
    • Experimental set-up: Field experiment
    • Experimental set-up: Greenhouse experiment
    • Statistical analysis
    • Data availability
  • Results
    • Field experiment
    • Greenhouse experiment
  • Discussion
    • Origin effects and maladaptation
    • The role of environmental conditions and soil-borne effects
  • Conclusion
  • Acknowledgements
  • References
  • Supplementary material
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