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  • Article title
  • Abstract
  • Keywords
  • Introduction
  • Methods
    • Study area and sampling
    • Chlorophyll content and fluorescence analysis
    • Analysis of soil physicochemical properties
    • Data handling and analysis
  • Results
  • Discussion
    • Non-invaded open dune vegetation
    • Invasion-induced changes in open dune vegetation and soil
    • Open dune vegetation response to the variability of invader thickets
  • Conclusions
  • Acknowledgements
  • References
  • Supplementary material
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